Education

BA, First Class Honors, University of Cambridge, England
PhD, University of Cambridge, England
Postdoctoral Fellow, The Salk Institute and University of Cambridge

Awards & Honors

• Szent-Györgyi Prize for Progress in Cancer Research, 2025
• Prince Mahidol Award (in Medicine), 2024
• American Association for Cancer Research Princess Takamatsu Memorial Lectureship Award, 2024
• AACR Lifetime Achievement Award in Cancer Research, 2022
• National Cancer Institute Outstanding Investigator, 2019
• Tang Prize for Biopharmaceutical Science, 2018
• Pezcoller-AACR International Award for Extraordinary Achievement in Cancer Research, 2018
• Sjöberg Prize for Cancer Research, 2017
• BBVA Foundation Frontiers of Knowledge Award, Biomedicine, 2015
• Royal Society, Royal Medal Award, 2014
• American Association for Cancer Research Fellow, 2013
• Clifford Prize for Cancer Research, 2007
• Robert J. and Claire Pasarow Award for Cancer Research, 2006
• Daniel Nathans Memorial Award, 2006
• American Philosophical Society Member, 2006
• Wolf Prize in Medicine, 2005
• Louisa Gross Horwitz Prize, 2004
• Prince of Asturias Award for Scientific and Technical Research, 2004
• Kirk A. Landon-AACR Prize for Basic Cancer Research, 2004
• American Cancer Society Medal of Honor, 2004
• National Academy of Medicine, 2004
• City of Medicine Award, 2003
• Sergio Lombroso Award in Cancer Research, 2003
• Keio Medical Science Prize, 2001
• J. Allyn Taylor International Prize in Medicine, 2000
• Gairdner Foundation International Award, 1994
• General Motors Cancer Research Foundation Prize, 1994
• American Academy of Arts and Sciences Fellow, 1992
• Royal Society of London Fellow, 1987
• American Cancer Society Research Professor
• United States National Academy of Sciences Member